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Mothers Against Decapentaplegic Homolog 9




  Name SMAD, Mothers against decapentaplegic homolog 9
  Gene HUGO code: SMAD9
  Type R-SMAD
  Functions Signal Transduction , Transcription
  Domains MH1 , MH2
  Taxa Phyla
  Cells many Heart , Brain , Placenta , Lung , Skeletal Muscle , Prostate , Testis , Ovary and Small Intestine
  Location Primary:
  Mods Phosphorylation
  Names '''SMAD 9, Mothers against DPP homolog 9, Smad9, Madh6'''


SMAD9 or Mothers against decapentaplegic homolog 9 is a polypeptide that, as its name describes, is a homolog of the ". It belongs to the SMAD family of proteins, which belong to the TGFβ superfamily of modulators. Like many other TGFβ family members, SMAD9 is involved in Cell Signalling . When a Bone Morphogenetic Protein binds to a receptor (BMP type 1 receptor kinase) it causes SMAD9 to interact with SMAD Anchor For Receptor Activation (SARA).The binding of ligands causes the Phosphorylation of the SMAD9 protein and the dissociation from SARA and the association with SMAD4 . It is subsequently transferred to the Nucleus where it forms complexes with other proteins and acts as a Transcription Factor . SMAD9 is a Receptor Regulated SMAD ( R-SMAD ) and is activated by bone morphogenetic protein type 1 receptor kinase. There are two Isoforms of the protein. Confusingly, it is also sometimes referred to as SMAD8 in the literature.